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Scientific Publications

To see the abstract of each article, roll your mouse over the authors' names (in red)
To see the articles themselves, click on the red text in each listing below.

If you have trouble accessing copy of articles, please email Adrian Treves


Treves, A. and M. Rabenhorst (0.21 Mb) (2017). Risk Map for Wolf Threats to Livestock still Predictive 5 Years after Construction. PLOS ONE: open access at

López-Bao*, J. V., G. Chapron and A. Treves* (2017).*equal co-authors (0.21 Mb) The Achilles heel of participatory conservation. Biological Conservation 212: 139–143: open access at DOI:

Treves, A., Artelle, K.A., Darimont, C.T., Parsons, D.R. 2017. (3.8 Mb) Mismeasured mortality: correcting estimates of wolf poaching in the United States. Journal of Mammalogy 98(3): open access at DOI:

Summary and FAQs (4 Mb)

Carroll, C., B. Hartl, G.T. Goldman, D.J. Rohlf, A. Treves, J.T. Kerr, E.G. Ritchie, R.T. Kingsford, K.E. Gibbs, M. Maron, J.E.M Watson. 2017.

Defending scientific integrity in conservation policy processes: lessons from Canada, Australia, and the United States.

Treves, A., Julia A. Langenberg, José v. López-Bao, Mark F. Rabenhorst 2017. (1.8 Mb)Gray wolf mortality patterns in Wisconsin from 1979 to 2012 Journal of Mammalogy 98(1): DOI:10.1093/jmammal/gyw145

Treves, A., Krofel, M., McManus, J. (equal co-authors).2016. Predator control should not be a shot in the dark. Frontiers in Ecology and the Environment14: 380-388. Click here to access the article at the journal website but don't forget the Supplement button up top.

Click here for short video explaining the findings

In a nutshell:

Treves, A., Bonacic, C. (equal co-authors). 2016. Humanity's Dual Response to Dogs and Wolves. Trends in Ecology and Evolution (TREE). doi:10.1016/j.tree.2016.04.006

Take-home message: The relationship between humans, dogs, and wolves has changed over more than 40,000 years in ways that reflect the ecology and evolved traits of all three species.

Chapron, G., Treves, A. (equal co-authors). 2016. Blood does not buy goodwill: allowing culling increases poaching of a large carnivore. Proceedings of the Royal Society B 283: 20152939.
Also see a recent update to our findings that make our conclusions even stronger.
Proceedings of the Royal Society B

Take-home messages: Culling wolves did not help Wisconsin or Michigan’s wolf populations. Culling did not reduce poaching and strongly appears to have increased poaching.

2015. Treves, A., Chapron, G., Lopez-Bao, J.V., Shoemaker, S., Goeckner, A., Bruskotter, J.T. Predators and the Public Trust. Biological Reviews doi: 10.1111/brv.12227

Take-home messages: Democratic governments have a duty under the public trust to preserve wildlife for current and future generations without substantial impairment. Trust duties are fiduciary duties meaning transparent accounting and prudence at a minimum. Few trustees of predaotrs have lived up these legal and ethical standards.

Treves, A., Krofel, M., López-Bao, J.V. Missing wolves, misguided policy. Science (eLetter) 350:1473-1475. Scroll to the bottom to see our concerns with Northern Rocky Mountain wolf mortality analyses.

Take-home message: Wolf-poaching was not adequately measured in the Northern Rocky Mountains population and governments did not intervene effectively or at all to prevent poaching.

Krofel, M., Treves, A., López-Bao, J.V. Hunted carnivores at outsized risk. Science. 350: 6260.

Take-home message: Large carnivores are particularly vulnerable to human-induced mortality and we still now little about how they respond..

Ripple, W. et al. 2016. Saving the World’s Terrestrial Megafauna. BioScience. doi: 10.1093/biosci/biw092.

2014. Treves, A. Bruskotter, J.T. Tolerance for Predatory Wildlife. Science 344: 476-477.

Take-home messages: Tolerance for predators did not reflect individual economic losses but rather social identity and peer group complaints. Intolerance for wolves and inclinations to poach wolves rose when the government culled wolves. Tolerance for wolves did not increase when wolf hunting and trapping season was implemented.

2015. Ohrens, O., Treves, A., Bonacic, C. Relationship between rural depopulation and puma-human conflict in the high Andes of Chile. Environmental Conservation doi:10.1017/S0376892915000259.

2015. Voyles, Z., Treves, A., Macfarland, D. Spatiotemporal effects of nuisance black bear management actions in Wisconsin. Ursus 26: 11-20.

2015. Hogberg, J., Treves, A., Shaw, B., Naughton-Treves, L. Changes in attitudes toward wolves before and after an inaugural public hunting and trapping season: early evidence from Wisconsin’s wolf range. Environmental Conservation, doi 10.1017/S037689291500017X.

2015. Browne-Nuñez, C., Treves, A., Macfarland, D., Voyles, Z., Turng, C. Tolerance of wolves in Wisconsin: A mixed-methods examination of policy effects on attitudes and behavioral inclinations. Biological Conservation 189: 59-71.


2014. Olson, E.R., Treves, A., Wydeven, A.P., Ventura, S. Landscape predictors of wolf attacks on bear-hunting dogs in Wisconsin, USA. Wildlife Research 41: 584–597.

2013. Bruskotter, J.T., Vucetich, J.A., Enzler, S., Treves, A., Nelson, M.P. Removing protections for wolves and the future of the U.S. Endangered Species Act (1973). Conservation Letters 7: 401-407.

2014. Bruskotter, J.T., Treves, A., Way J.G. Carnivore Management. pp. 83-90 in B.S. Steel editor. Science and Politics: An A-To-Z Guide to Issues and Controversies. CQ Press, Thousand Oaks, CA, USA. .

2013. Treves, A., Naughton-Treves, L., Shelley, V. Longitudinal Analysis of Attitudes Toward Wolves. Conservation Biology 27: 315–323

2012. Treves, A. Tolerant attitudes reflect an intent to steward: A Reply to Bruskotter and Fulton. Society and Natural Resources 25: 103-104.

2011. Shelley, V., Treves, A., Naughton, L. Attitudes to Wolves and Wolf Policy Among Ojibwe Tribal Members and Non-tribal Residents of Wisconsin's Wolf Range. Human Dimensions of Wildlife 16: 397-413.

2011. Treves, A., Bruskotter, J. Gray Wolf Conservation at a Crossroads. BioScience 61: 584-585.
2011. Bruskotter, J., Enzler, S., Treves, A. Rescuing Wolves from Politics: Wildlife as a Public Trust Resource. Science (Wash., D.C.), Policy Forum 333(6051): 1828-1829.

2011. Treves, A., Martin K.A. Hunters as stewards of wolves in Wisconsin and the Northern Rocky Mountains, USA. Society and Natural Resources 24: 984-994.

2011. Treves, A., Martin K.A., Wydeven, A.P., Wiedenhoeft, J.E. Forecasting Environmental Hazards and the Application of Risk Maps to Predator Attacks on Livestock. Bioscience 61(6): 451-458.

2011. Schloegel, C., Jones, T., Zug, B., Achig, L., Treves, A. Don Oso Program Develops Participatory Monitoring Protocol for Andean Bears in Southern Sangay National Park, Ecuador. International Bear News 20(2): 23-25.

2010. Treves, A., Jones, S.M. Strategic trade-offs for wildlife-friendly eco-labels. Frontiers in Ecology and the Environment 8(9): 491–498.

2010. Treves, A., Kapp, K.J., Macfarland, D.M. American black bear nuisance complaints and hunter take. Ursus 21(1):30-42.

2010. Treves, A., Mwima, P., Plumptre, A.J., Isoke, S. Camera-trapping forest-woodland wildlife of western Uganda reveals how gregariousness biases estimates of relative abundance and distribution. Biological Conservation. 143: 521-528.

2009. Treves, A. Hunting for large carnivore conservation. Journal of Applied Ecology 46: 1350-1356.

2009 Treves, A., Jurewicz, R., Naughton-Treves, L., Wilcove, D. The price of tolerance: wolf damage payments after recovery. Biodiversity and Conservation, 2009, 18(14):4003-4021.

2009. Treves, A., Wallace, R.B., White, S. Participatory planning of interventions to mitigate human-wildlife conflicts. Conservation Biology 2009, 23(6): 1577-1587.

2009. Treves, A., Plumptre, A.J., Hunter, L.T.B., Ziwa, J. Identifying a potential lion Panthera leo stronghold in Queen Elizabeth National Park, Uganda, and Parc National des Virunga, Democratic Republic of Congo. Oryx 43(1): 60-66. (Copyright Cambridge University Press and see the Erratum we published as a correction to this manuscript.

2008. Treves, A. Beyond Recovery: Wisconsin's Wolf Policy 1980-2008. Human Dimensions of Wildlife 13(5): 329-338.

2008. Treves, A. Human-wildlife conflicts around protected areas. In Wildlife and Society: The Science of Human Dimensions. Manfredo, M., Vaske, J. J., Brown, P., Decker, D.J., Duke, E.A. eds. Island press, Washington, DC.

2009. Treves, A., Martin, K.A., Wiedenhoeft, J.E., Wydeven, A.P. Gray wolf dispersal in the Great Lakes Region, in Recovery of Gray Wolves in the Great Lakes Region of the United States: an Endangered Species Success Story. Wydeven, A. P., Heske, E.H., Van Deelen, T. R. eds. Springer: New York.

2007. Treves, A., Palmqvist, P. Reconstructing hominin interactions with mammalian carnivores (6.0 - 1.8 Ma) in Primate Anti-Predator Strategies. K. A. I. Nekaris, and S. L. Gursky, eds. Springer, New York.

2007. Sillero-Zubiri, C., Sukumar, R., Treves, A. Living with wildlife: the roots of conflict and the solutions. In: Key Topics in Conservation Biology. D. MacDonald and K. Service, eds. Oxford, Oxford University Press, pp. 266-272.

2006. Treves, A., Wallace, R.B., Naughton-Treves, L., Morales, A. Co-managing human-wildlife conflicts: A review. Human Dimensions of Wildlife 11(6):1-14

Or you can read the Policy Brief of this article.

2006. Treves, A., Andriamampianina, L., Didier, K., Gibson, J,. Plumptre, A., Wilkie, D., Zahler, P. A simple, cost-effective method for involving stakeholders in spatial assessments of threats to biodiversity. Human Dimensions of Wildlife 11(1): 43-54.

2005. Treves, A., Naughton-Treves, L. Evaluating lethal control in the management of human-wildlife conflict. People and Wildlife, Conflict or Coexistence? Woodroffe, R., Thirgood, S., Rabinowitz, A. eds. Cambridge University Press, Cambridge, UK. pp. 86-106.

2005. Treves, A., Brandon, K. Tourism impacts on the behavior of black howler monkeys (Alouatta pigra) at Lamanai, Belize. In Commensalism and Conflict: The primate-human interface. J. Paterson & J. Wallis, eds. American Society of Primatology, Norman, OK.

2005. Naughton-Treves, L., Treves, A. Socioecological factors shaping local tolerance of crop loss to wildlife in Africa. People and Wildlife, Conflict or Coexistence? Woodroffe, R., Thirgood, S., Rabinowitz, A. eds. Cambridge Univ. Press, Cambridge, UK. pp. 253-277.

2004. Treves, A., Naughton-Treves, L. Harper, E., Mladenoff, D., Rose, R., Sickley, T., Wydeven, A. Predicting human-carnivore conflict: A spatial model derived from 25 years of wolf predation on livestock. Conservation Biology 18(1): 114-125.

2004. Wydeven, A.P., Treves, A., Brost, B., Wiedenhoeft, J. Characteristics of wolf packs in Wisconsin: Identification of traits influencing depredation. Pp. 28-50 in People and Predators: From Conflict to Coexistence. N. Fascione, A. Delach, M. Smith, eds. Island Press, Washington, DC

2003. Treves, A., Karanth, K.U. Human-carnivore conflict and perspectives on carnivore management worldwide. Conservation Biology 17(6): 1491-1499.

2003. Grossberg, R., Treves, A., Naughton-Treves, L. Incidental ecotourism at Lamanai, Belize: The incidental ecotourist - Measuring visitor impacts on endangered howler monkeys inhabiting an archaeological site in Belize. Environmental Conservation 30(1): 40-51.

2003. Naughton-Treves, L., Grossberg, R., Treves, A. Paying for tolerance: The impact of livestock depredation and compensation payments on rural citizens' attitudes toward wolves. Conservation Biology 17(6): 1500-1511.

2003. Naughton-Treves, L., Mena, J.L., Treves, A., Alvarez, N., Radeloff, V.C. Wildlife survival beyond park boundaries: The impact of swidden agriculture and hunting on mammals in Tambopata, Peru. Conservation Biology 17(4): 1106-1117.

2003. Shivik, J.A., Treves, A., Callahan, M. Nonlethal techniques for managing predation: primary and secondary repellents. Conservation Biology 17(6): 1531-1537.

2002. Treves, A., Jurewicz, R., Naughton-Treves, L., Rose, R., Willging, R., Wydeven, A. Wolf depredation on domestic animals in Wisconsin, 1976-2000. Wildlife Society Bulletin 30:231-241.

Pre-wolf research

1999. Treves, A., Naughton-Treves, L. Risk and opportunity for humans coexisting with large carnivores. Journal of Human Evolution 36: 275-282.

1998. Naughton-Treves, L., Treves, A., Chapman, C., Wrangham, R.W. Temporal patterns of crop raiding by primates: Linking food availability in croplands and adjacent forest. Journal of Applied Ecology 35(4): 596-606.

For earlier articles about primate behavioral ecology, please email Adrian Treves.