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Adrian Treves

Professor and Founder of the Carnivore Coexistence Lab

Adrian Treves conducts independent research and advocates for future generations of all life, for scientific integrity, and for sovereign publics worldwide. He studies and speaks about the public trust doctrine and intergenerational equity around the world More on public trust doctrines and intergenerational equity here. Adrian earned his PhD at Harvard University in 1997 and is a Professor of Environmental Studies at the University of Wisconsin–Madison and director of the Carnivore Coexistence Lab since 2007. For the past 27 years, his research focuses on ecology, law, and human dimensions of ecosystems in which crop and livestock ownership overlaps the habitat of large carnivores from coyotes up to grizzly bears. He has authored >133 scientific papers on predator-prey ecology or conservation.
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30A Science Hall
550 N. Park Street
Madison, WI 53706
Phone: (608) 890-1450
Email: atreves [at]

Google Scholar profile here

The University of Wisconsin–Madison occupies ancestral Ho-Chunk land, a place their nation has called Teejop (day-JOPE) since time immemorial. In an 1832 treaty, the Ho-Chunk were forced to cede this territory. Decades of ethnic cleansing followed when both the federal and state government repeatedly, but unsuccessfully, sought to forcibly remove the Ho-Chunk from Wisconsin. This history of colonization informs our shared future of collaboration and innovation.Today, UW–Madison respects the inherent sovereignty of the Ho-Chunk Nation, along with the eleven other First Nations of Wisconsin.

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Post docs

“Francisco” Francisco Santiago-Avila, MPP, MEM (Ph.D. 2019)

Fran holds Masters degrees in environmental public policy as well as environmental management from Duke University. His recent research has focused on valuation of non-wood forest wealth, the prioritization of ecosystems for conservation in Latin America and the Caribbean, and measuring the impact of protected areas on land cover change. As part of CCL, Fran's research has revolved around the integration and application of environmental and animal ethics to coexistence with wildlife, and the evaluation of the effectiveness of lethal and non-lethal methods to prevent conflicts with large carnivores (the gray wolf, in particular). His main objective is to reform wildlife management by embedding in it the much needed acknowledgement of moral standing for individual nonhuman animals. Fran's other research interests include: human behavior and attitudes towards animals, conflict-mitigating interventions and legal mechanisms for conservation. His other interests include carnivore behavioral ecology and trophic cascades

Treves, A., Santiago-Ávila, F., Lynn, W.S. (equal co-authors) 2018. Just Preservation. Biological Conservation 229: 134-141.

Santiago-Avila, F.J., Cornman, A.M., Treves, A. 2018. Killing wolves to prevent predation on livestock may protect one farm but harm neighbors. PLOS One. 10.1371/journal.pone.0189729

2019. Ohrens, O., Santiago-Avila, F.J., Treves, A. The Twin Challenges of Preventing Real and Perceived Threats to Human Interests. Chapter 12 in: Human-Wildlife Interactions: Turning Conflict into Coexistence. Eds. Frank, B., Marchini, S., Glikman, J. Cambridge University Press.

Graduate Students

“Drew” Drew Bantlin, M.S. (Ph.D. candidate)

Drew earned his M.S. in 2017 in Environment and Resources from UW-Madison. Drew's research focuses on the effect of African lion reintroduction to Akagera National Park, Rwanda. He is exploring the trophic cascade hypothesis, that top carnivores have top-down influence on other species in an ecosystem. He is examining how lions may influence the behavior of other species in the park, which may in turn have effects on plant communities and ecosystem processes. Much of his field work involves direct observations of the lions and their prey, camera trapping, and analysis of GPS collared animals. In addition to research, Drew also assists Akagera with other research projects, including assisting with monitoring the newly reintroduced black rhinos, conducting population surveys on the ground and in the air, and collecting data on the demographics of elephants, giraffes, hyenas, and leopards. Prior to coming to Akagera, Drew has had field experience working with gray wolves, mountain gorillas, chimpanzees, and numerous monkey species.

Bantlin, Drew A. 2018.Reintroduction of African Lions to Akagera National Park, Rwanda. Global Reintroduction Perspectives, 2018: Case Studies from around the Globe, by Pritpal S. Soorae, 6th ed., IUCN/SSC Reintroduction Specialist Group, 2018, pp. 182–186.

“Karann” Karann Putrevu (Ph.D. candidate)

Karann is a computational biologist originally from St. Catharines, Ontario, Canada. He completed his Bachelor of Science with a dual degree in biological engineering and biology at Cornell University. He has a particularly strong passion for tigers and has studied topics ranging from modeling their interspecific conflicts (with leopards in India and wolves in Russia) to the Panthera genus' genetic lineage to the social and political context of the South China tiger's extinction in the wild. Karann is currently working on studying the predator ecology of the Amur tiger in the Russian Far East, applying Bayesian methods to model large carnivore population dynamics, and modeling spatial risk of poaching.

“Naomi” Naomi Louchouarn, MESM (Ph.D. candidate)

Naomi graduated from McGill University in 2014 with a Bachelor’s of Science in Environmental Biology and dual minors in wildlife biology and applied ecology. During and after her Bachelor’s degree Naomi worked on various research projects in Northern Canada, South Africa and Australia. These experiences provided her with the inspiration to pursue research focused on conservation planning and mitigating conflicts between humans and wildlife. Naomi received a Master’s of Environmental Science and Management at UC-Santa Barbara’s Bren School in 2017. Her Master’s research focused on prioritizing land for conservation in Canada’s Mackenzie River Basin to maximize the protection of caribou habitat and minimize economic losses to resource extraction industries in the region. With the CCL, Naomi will lead a project focused on identifying non-economic motivations for poaching of large carnivores (such as grizzly bears) in the North American West. The objective of this project will be to model when and where large carnivore poaching is likely to occur. She is also leading a risk mapping effort to predict where livestock losses to wolves are likely to occur.


Alexandra Pineda Guerrero, Professional Masters in Environmental Conservation (Ph.D. candidate)

Alexandra is a biologist from Bogotá, Colombia and received a Master’s degree of Environmental Conservation at UW-Madison in 2016. She has worked as a researcher in the public and private sector in Colombia. Her work has been related to the ecology of carnivores, conservation planning for tropical mammals and ecosystems, and human-carnivore coexistence and conflicts, specifically related to jaguar and pumas. Alexandra won a scholarship from the Colombian government – COLCIENCIAS- for her Ph.D and she is interested in testing the efficacy of non-lethal methods of preventing jaguar and puma attacks in Colombia.


“Brian” Brian Schuh, Professional Masters in Environmental Conservation

Brian is a wildlife conservationist originally from Santa Barbara, California, who recently completed his Master's degree from the University of Wisconsin-Madison. During his bachelor's degrees in Zoology and Botany, also from UW-Madison, he worked on conservation projects with lumholtz tree kangaroos in Australia, whale sharks in Mexico, and cheetahs in South Africa. These experiences cemented in him a passion for wildlife conservation, particularly focusing on international human-wildlife coexistence with a special interest in cheetahs. During his Master's project he partnered with the Kenya Wildlife Trust (KWT) working to mitigate human-wildlife conflict and understand the population dynamics of lions and cheetahs in the Maasai Mara. Continuing this partnership with KWT, his current research aims at identifying human and natural variables affecting the distribution and abundance of cheetahs in the Maasai Mara and developing management srecommendations for their long term survival.

Grad Packet for use within the UW G suite, find a living document you can edit as we learn together about the graduate career.


Annilee Kremling, Natalia Vonarburg, Kaari Hostler (the Constitution Project)

Affiliates at other universities

“Camilla” Camilla Fox, M.A.

Camilla H. Fox is the founder and executive director of Project Coyote- a national non-profit organization that promotes coexistence between people and wildlife through education, science, and advocacy. With 25+ years of experience working on behalf of wildlife and wildlands and a Masters degree on wildlife ecology, policy, and conservation, Camilla’s work has been featured in several films, books and national media outlets. A frequent speaker on these issues, Camilla is co-author of two books— Coyotes in Our Midst and Cull of the Wild and is co-producer of the award-winning documentary Cull of the Wild ~ The Truth Behind Trapping. She produced and directed the award-winning documentary film KILLING GAMES ~ Wildlife in the Crosshairs, released in 2017. In 2006, Camilla received the Humanitarian of the Year Award from the Marin Humane Society and the Christine Stevens Wildlife Award from the Animal Welfare Institute. She was named one of the 100 Guardian Angels of the Planet in 2013 and the 2014 Conservationist of the Year Award by the John Muir Association. In 2016 she was honored with the Grassroots Activist of the Year Award by the Fund for Wild Nature. See the PC webpage.

“Suzanne” Dr. Suzanne Agan (Ph.D. from Antioch University of New England)

Suzanne is working on the human dimensions and spatial ecology of poaching and implications for red wolf survival in North Carolina.

bioRxiv pre-publication manuscript on red wolf mortality risk.

“Jeannine” Jeannine McManus, Ph.D.

Jeannine is interested in the scientific underpinning of practical conservation biology solutions to major conservation challenges. Jeannine has 10 years of conservation work experience which has focused on human-wildlife conflict resolution by linking practical and economical tools; biodiversity conservation, within and outside of protected areas; and environmental education. Jeannine ha worked on research projects comparing the economic and ecological effects of production landscapes on biodiversity; meta-population structure, spatial ecology, behaviour, and resource selection of leopard (Panthera pardus) in the Eastern and Western Cape, South Africa. Jeannine passed my PhD in Ecology at Witwatersrand University, and I am currently working on an arid region restoration project in the Karoo, South Africa with the Landmark Foundation.

Tshabalala, T., McManus, J., Treves, A., Masocha, V., Faulconbridge, S., Schurch, M., Goets, S., Smuts, B. 2021. Leopards and mesopredators as indicators of mammalian species richness across diverse landscapes of South Africa. Ecological Indicators 121, 107201.

no photo found Arlyne Johnson, Ph.D.

Arlyne is an Honorary Fellow with the Nelson Institute and works with Foundations of Success providing training for conservation organizations in the U.S. and abroad in designing and monitoring the effectiveness of conservation projects. She has led wildlife research and conservation programs in Latin American and Asia for the Wildlife Conservation Society since 1988, including projects to reduce conflict between local communities and large carnivores (tiger, dhole) and Asian elephant in Lao PDR. Arlyne currently teaches a graduate-level course in Conservation Planning for the Nelson Institute. She holds a Ph.D. in Environment and Resources from the Nelson Institute.

Jaimie in Bridger Jamie Hogberg, M.S.

Jamie received her MS in Conservation Biology and Sustainable Development at UW-Madison, where she studied human-wolf conflict in Wisconsin and attitudes toward wolves and management following the inaugural public hunting and trapping season. She has worked as a field biologist for conservation organizations in the western U.S., tropical conservation in South America, and she currently coordinates the Environmental Conservation MS Program at the Nelson Institute. Jamie is also the founding board member of the Wisconsin Chapter of the Society for Conservation Biology. Her interests include human dimensions of wildlife conservation, private lands conservation, avian ecology, and community-based conservation planning.

Nathan Schulfer, M.S.

Nathan began work for the Environmental Conservation Professional Masters program at the Nelson Institute after graduating in 2014 with a Masters degree on food security and ecosystem services, co-advised by Dr. A. Treves and Dr. J. Silbernagel.

Congratulations to graduates 2012–present

Abigail M. Fergus, MS, 2020

Francisco Santiago-Ávila, Ph.D. 2019

Omar Ohrens, M.S. 2013, Ph.D. 2018

Drew Bantlin, M.S. 2016

Jamie Hogberg, M.S. 2014

Zachary Voyles, M.S. 2013

Brittany Bovard, M.S. 2013

Professional Masters in Environmental Conservation 2012–present

Brian Schuh, 2018

Elena Jove-Edens, 2017

Alexandra Pineda Guerrero, 2016

Naseem Sultani, 2015

Paula Henriquez, 2014

Non-thesis Masters in Conservation Biology and Sustainable Development

Doris Ming Hua, M.S. 2014

Nathan Schulfer, M.S. 2012

For older records, email Adrian Treves